Chester, a popular gray mouse who lives on Anastasia Island, in St. Augustine, Florida discovers that four of his friends have accidentally been taken to Savannah, Georgia. They were in the back of a neighbor’s old car that was being hauled there for repairs. Mr. Dalton and Chester travel to Savannah to find them and bring them home. While there, Mr. Dalton has a truck accident that leads them to meeting friends of Fletcher and Henrietta.